Morley Read / Leaf frog (Agalychnis hulli) blinking its eyes and turning around, Orellana Province, Ecuador. (non-ex)
01446163
Ingo Arndt / Red-eyed tree frog (Agalychnis callidryas) eye closed with semi-transparent eyelid that allows it to see its surroundings even while resting, Costa Rica.
01446162
Ingo Arndt / Red-eyed Tree Frog (Agalychnis callidryas) close- up of eye half open, the golden webbed semi-transparent eyelid allows to see its surrounding even while resting, Costa Rica.
01516488
Richard Robinson / A Blue shark (Prionace glauca) just below the surface, with nictitating membrane partially shut to protect its eye, Hauraki Gulf, Auckland, New Zealand, June.
01443744
Bence Mate / Dalmatian pelicans (Pelecanus crispus) underwater view of group feeding, Lake Kirkini, Greece. Highly commended in the GDT competition 2013. Commended in the Wildlife Photographer of the Year Awards c...
Jack Dykinga / Raven (Corvus corax) with nictitating membrane over eye, in a blizzard on Hurricane Ridge, Olympic National Park, Washington, USA. Sequence 2/2
01606675
John Waters / Common European toad (Bufo bufo) at surface, retreats underwater, Priddy, Somerset, England, UK, April.
01601156
Eric Baccega / Shoebill stork (Balaeniceps rex) portrait with a closed eyelid. Swamps of Mabamba, Lake Victoria, Uganda.
01594249
Jordi Chias / Whale shark (Rhincodon typus), closing nictitating membrane, South Ari Atoll, Maldives Islands, Indian Ocean.
01590286
Nick Upton / Rook (Corvus frugilegus) calling while perched in a Willow tree and blinking to show its blue nictitating membrane, Gloucestershire, UK, February.
01535401
Dave Bevan / European common frog (Rana temporaria) in a pond, closing its nictating membranes to lubricate its eyes, Carmarthenshire, Wales, UK, February.
01446168
Ingo Arndt / Spurell's Flying Frog (Agalychnis spurelli) eye closed with semi-transparent eyelid that allows it to see its surroundings even while resting, Costa Rica.
01526207
Juan Carlos Munoz / Bateleur eagle (Terathopius ecaudatus) portrait with nictitating membrane closed, Captive, occurs in Africa.
01516490
Richard Robinson / Blue shark (Prionace glauca) with nictitating membrane partially shut to protect its eye, Hauraki Gulf, Auckland, New Zealand, June.
01516489
Richard Robinson / Blue shark (Prionace glauca) with mouth open and nictitating membrane almost completely shut to protect its eye, Hauraki Gulf, Auckland, New Zealand, June.
01516487
Richard Robinson / A Blue shark (Prionace glauca) just below the surface, with nictitating membrane partially shut to protect its eye, Hauraki Gulf, Auckland, New Zealand, June.
01477285
David Tipling / Rook (Corvus frugilegus) calling, Norfolk, February.
01447814
Chris Mattison / Red-eyed leaf frog (Agalychnis callidryas) close-up of eye showing patterned lower eyelid.
01445350
Roland Seitre / Bocourt's Eyelid Skink (Phoboscincus garnieri) captive, Parc zoologique et forestier / Zoological and Forest Park, Noumea, South Province, New Caledonia
01442514
Dave Watts / Bateleur (Terathopius ecaudatus) showing nictitating membrane. Kgalagadi National Park, South Africa
01442512
Dave Watts / Bateleur (Terathopius ecaudatus) showing nictitating membrane. Kgalagadi National Park, South Africa
00499151
Ingo Arndt / Gliding Leaf Frog (Agalychnis spurrelli) eye closed with semi-transparent eyelid that allows it to see its surroundings even while resting, Costa Rica
00499149
Ingo Arndt / Red-eyed Tree Frog (Agalychnis callidryas) eye closed with semi-transparent eyelid that allows it to see its surroundings even while resting, Costa Rica
00499139
Ingo Arndt / Red-eyed Tree Frog (Agalychnis callidryas) semi-transparent eyelid that allows it to see its surroundings even while resting, Costa Rica
00482804
Thomas Marent / Red-eyed Tree Frog (Agalychnis callidryas) with half closed eyelid, Tortuguero National Park, Costa Rica
01298973
Jeff Rotman / Eye of Tiger shark (Galeocerdo cuvier), nictitating membrane closing to protect eye, Great Barrier Reef, Australia, Coral Sea.
01257243
Mark Bowler / Red eyed tree frog (Agalychnis callidryas) showing two of its three eyelids, patterned eyelid completely closed over left eye Captive
01233224
Wild Wonders of Europe / Varesvuo / European Roller (Coracias garrulus) preening, Pusztaszer, Hungary, May 2008
01185770
Doug Perrine / Tiger shark (Galeocerdo cuvier) lunging for bait, showing nictitating membrane covering eye to protect it, North Shore, Oahu, Hawaii, USA, Central Pacific Ocean
01185769
Doug Perrine / Tiger shark (Galeocerdo cuvier) lunging for bait, showing nictitating membrane covering eye to protect it, North Shore, Oahu, Hawaii, USA, Central Pacific Ocean
01181522
Philippe Clement / Arizona toad (Bufo microscaphus) burrowing in mud and showing the transparent inner eyelid or nictating membrane, Arizona, USA. Captive
01181523
Philippe Clement / Arizona toad (Bufo microscaphus) burrowing in mud and showing the transparent inner eyelid or nictating membrane, Arizona, USA. Captive
01171439
Jane Burton / Border Collie wearing Elizabethan collar to prevent scratching, after operation to stitch lower eyelid
01171450
Jane Burton / Border Collie, 12-year, in Elizabethan collar to prevent scratching of sutured lower eyelid
00298284
Piotr Naskrecki / Fantastic Leaf-tail Gecko (Uroplatus phantasticus) cleaning eyelid's transparent shield with its large, muscular tongue, Madagascar
00298104
Piotr Naskrecki / Red-eyed Tree Frog (Agalychnis callidryas) semi-transparent eyelid that allows it to see its surroundings even while resting, Costa Rica
01115267
John Cancalosi / Regal horned lizard discharging blood from eyelid in defense {Phrynosoma solare}
00499148
Ingo Arndt / Red-eyed Tree Frog (Agalychnis callidryas) eye half covered with semi-transparent eyelid that allows it to see its surroundings even while resting, Costa Rica
01739274
Joel Sartore / Photo Ark / Violet-necked lory (Eos squamata obiensis) looking at the camera and vocalising, Anak Burung Tropikana, Bali. Captive.
01739273
Joel Sartore / Photo Ark / Violet-necked lory (Eos squamata obiensis) looking at the camera and blinking, Anak Burung Tropikana, Bali. Captive.
01739279
Joel Sartore / Photo Ark / Oriental honey-buzzard (Pernis ptilorhynchus orientalis) looking around and blinking, Bali Bird Park, Bali. Captive.
01728832
Klaus Nigge / Philippine eagle (Pithecophaga jefferyi) blinking with crest raised, showing nictitating membrane, Philippine Eagle Centre, Malagos, Davao, Mindanao, Philippines, August 2006. Critically endangered....
01730441
Andy Rouse / Brown fish owl (Ketoupa zeylonensis) fishing in shallow water, with nictitating membrane visible on one eye, Ranthambhore, India.
01730439
Andy Rouse / Brown fish owl (Ketoupa zeylonensis) spreading wings whilst fishing in shallow water, Ranthambhore, India.
01726733
John Cancalosi / Timelapse of Common snapping turtle (Chelydra serpentina) lifting head out of muddy water, blinking and then submerging again, Maryland, USA.
01725088
Eric Baccega / Buzzard (Buteo buteo) with nictitating membrane visible, perched on tree stump, Lorraine, France. March.
01723248
Joel Sartore / Photo Ark / Iraqi eyelid gecko (Eublepharis angramainyu) head portrait, Saint Louis Zoo. Captive, occurs in Middle East.
01723245
Joel Sartore / Photo Ark / Iraqi eyelid gecko (Eublepharis angramainyu) portrait, Saint Louis Zoo. Captive, occurs in Middle East.
01723244
Joel Sartore / Photo Ark / Iraqi eyelid gecko (Eublepharis angramainyu) portrait, Saint Louis Zoo. Captive, occurs in Middle East.
01719474
Daniel Nunez / Red eyed tree frog (Agalychnis callidryas) pair in amplexus at night, Calakmul Biosphere Reserve, Campeche, Mexico. Cropped.
01719465
Daniel Nunez / Black-eyed tree frogs (Agalychnis moreleti) pair in amplexus, male with nictitating membrane visible, Finca El Amate, Escuintla, Guatemala.
01719464
Daniel Nunez / Black-eyed tree frogs (Agalychnis moreleti) pair in amplexus, male with nictitating membrane visible, Finca El Amate, Escuintla, Guatemala.
01715164
Manuel Gracia / Booted eagle (Hieraaetus pennatus) looking around and calling. The animal blinks showing its nictitating membrane. Seville, Spain.
01715158
Manuel Gracia / Booted eagle (Hieraaetus pennatus) head-shot. The animal blinks showing its nictitating membrane. Seville, Spain.
01715157
Manuel Gracia / Tracking shot of a Booted eagle (Hieraaetus pennatus) standing on a log and cleaning its beak. The animal blinks repeatedly showing its nictitating membrane. Seville, Spain.
01714460
Joel Sartore / Photo Ark / Starrett's frog (Isthmohyla tica) profile. The animal seated, breathing. The animal then closes its eye, showing its nictitating membrane. La Libia, Costa Rica. Critically Endangered. Controlled condi...
01714375
Joel Sartore / Photo Ark / A close up of a Brolga crane's (Grus rubicunda) head. The animal blinks repeatedly and looks around. International Crane Foundation, Baraboo, Wisconsin, USA. Captive.
01714369
Joel Sartore / Photo Ark / Mountain yellow-legged frog (Rana muscosa) lying down, breathing. The animal opens its eye, showing its nictitating membrane. Then the frog jumps out of the frame. Oakland Zoo, California, USA. Captiv...
01714368
Joel Sartore / Photo Ark / Mountain yellow-legged frog (Rana muscosa) lying down and opening its eye, showing its nictitating membrane. The animal is showing respiration. The frog then jumps out of the frame. Oakland Zoo, Calif...
01714367
Joel Sartore / Photo Ark / Foothill yellow-legged frog (Rana boylii), sitting upright, breathing. The animal is opening and closing its eye, showing its nictitating membrane. Oakland Zoo, California, USA. Captive.
01714440
Joel Sartore / Photo Ark / Egyptian fruit bat (Rousettus aegyptiacus) lying down and looking at the camera. The animal blinks. Omaha?s Henry Doorly Zoo and Aquarium, Nebraska, USA. Captive.
01714430
Joel Sartore / Photo Ark / A close up of a Yellow-bellied marmot (Marmota flaviventris) juvenile male's head looking at the camera. The animal blinks. Calgary Wildlife, Canada. Captive.
01714396
Joel Sartore / Photo Ark / A close up of a Bulwer?s pheasant's (Lophura bulweri) head. The animal repeatedly blinks and then looks at the camera. This animal is the last known female in captivity. Al Bustan Zoological Centre, S...
01714394
Joel Sartore / Photo Ark / Senegal dove (Streptopelia senegalensis) looking around and blinking, Lincoln Children's Zoo, Nebraska, USA. Captive.
01708191
Franco Banfi / Alpine newt (Ichthyosaura alpestris) swimming in lake, nictitating membrane visible, portrait, upper Verzasca valley, Ticino, Switzerland. August.
01713181
Harry Collins / Belted kingfisher (Megaceryle alcyon) perched overlooking a small pond. Animal is showing nictitating membrane. Florida, USA.
01713185
Manuel Gracia / Ocellated lizard (Timon lepidus) sitting on a rock and basking in the sun, Sevilla, Sierra Morena, Spain.
01712578
Harry Collins / African leopard (Panthera pardus pardus) lying in the shade. Animal blinks and its ears move as it listens. Mara Triangle, Maasai Mara National Reserve, Kenya, Africa, November.
01710192
Neil Aldridge / Long-nosed horned frog (Megophrys nasuta) close up of eye and elongated upper eyelid (horns), Danum Valley, Sabah, Borneo.
01711405
Neil Aldridge / Common toad (Bufo bufo) in a meadow closing and opening eye showing nictitating membrane and eyelid, Bunloit Estate, Scotland, summer.
01709262
Harry Collins / Peregrine falcon (Falco peregrinus) perched and closing eye showing nictitating membrane and eyelid, New Jersey, USA, June.
01705551
Joel Sartore / Photo Ark / White-tailed mongoose (Ichneumia albicauda) looking around, Arabia's Wildlife Centre. Captive.
Joel Sartore / Photo Ark / Black francolin (Francolinus henrici) male close up of head looking around, Wadi Al Safa Wildlife Center. Captive.
01705486
Joel Sartore / Photo Ark / Egyptian turtle dove (Streptopelia turtur rufescens) looking around, Sheikh Butti Maktoum Wildlife. Captive.
01705485
Joel Sartore / Photo Ark / Arabian eagle owl (Bubo milesi) female close up of head looking around, Kalba Bird of Prey Center. Captive.
01705484
Joel Sartore / Photo Ark / Arabian eagle owl (Bubo milesi) female close up of head looking around, Kalba Bird of Prey Center. Captive.
01705483
Joel Sartore / Photo Ark / Mottled owl (Ciccaba virgata centralis) close up of head looking around, Toucan Rescue Ranch. Captive.
01705482
Joel Sartore / Photo Ark / Mottled owl (Ciccaba virgata centralis) close up of head looking around, Toucan Rescue Ranch. Captive.
01705481
Joel Sartore / Photo Ark / Black-and-white owl (Ciccaba nigrolineata) close up of head looking around, Toucan Rescue Ranch. Captive.
01705474
Joel Sartore / Photo Ark / Northern blue jay (Cyanocitta cristata bromia) looking around, profile. Captive.
01708802
Roland Seitre / King vulture (Sarcoramphus papa) yawning, showing its protective nictitating membrane or "third eyelid", head portrait, Parque Jaime Duque, Bogota, Colombia. Captive.
01706930
David Fleetham / Galapagos shark (Carcharhinus galapagensis), listed as potentially dangerous, at sea surface, with nictitating membrane covering its eye, Hawaii, Pacific Ocean.
01704847
David Weiller / Long-nosed horned frog (Pelobatrachus nasutus) juvenile close up portrait, closing its eyes, showing the nictitating membrane, Crocker Range, Sabah, Borneo, July.
01702441
Joel Sartore / Photo Ark / European Tengmalm's owl (Aegolius funereus) close up of face, International Centre for Birds of Prey. Captive.
01700495
Joel Sartore / Photo Ark / Elf owl (Micrathene whitneyi) mid shot portrait of head, blinking. Captive.