Jack Dykinga / Raven (Corvus corax) with nictitating membrane over eye, in a blizzard on Hurricane Ridge, Olympic National Park, Washington, USA. Sequence 2/2
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John Waters / Common European toad (Bufo bufo) at surface, retreats underwater, Priddy, Somerset, England, UK, April.
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Eric Baccega / Shoebill stork (Balaeniceps rex) portrait with a closed eyelid. Swamps of Mabamba, Lake Victoria, Uganda.
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Jordi Chias / Whale shark (Rhincodon typus), closing nictitating membrane, South Ari Atoll, Maldives Islands, Indian Ocean.
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Nick Upton / Rook (Corvus frugilegus) calling while perched in a Willow tree and blinking to show its blue nictitating membrane, Gloucestershire, UK, February.
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Dave Bevan / European common frog (Rana temporaria) in a pond, closing its nictating membranes to lubricate its eyes, Carmarthenshire, Wales, UK, February.
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Ingo Arndt / Spurell's Flying Frog (Agalychnis spurelli) eye closed with semi-transparent eyelid that allows it to see its surroundings even while resting, Costa Rica.
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Juan Carlos Munoz / Bateleur eagle (Terathopius ecaudatus) portrait with nictitating membrane closed, Captive, occurs in Africa.
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Richard Robinson / Blue shark (Prionace glauca) with nictitating membrane partially shut to protect its eye, Hauraki Gulf, Auckland, New Zealand, June.